PUBLICATION

Structures of CYLD USP with Met1- or Lys63-linked diubiquitin reveal mechanisms for dual specificity

Authors
Sato, Y., Goto, E., Shibata, Y., Kubota, Y., Yamagata, A., Goto-Ito, S., Kubota, K., Inoue, J.I., Takekawa, M., Tokunaga, F., Fukai, S.
ID
ZDB-PUB-150217-1
Date
2015
Source
Nature structural & molecular biology   22(3): 222-9 (Journal)
Registered Authors
Keywords
none
MeSH Terms
  • Ubiquitin Thiolesterase/chemistry
  • HEK293 Cells
  • Crystallography, X-Ray
  • Protein Structure, Tertiary
  • Conserved Sequence
  • Humans
  • Kinetics
  • Models, Molecular
  • Zebrafish Proteins/chemistry*
  • Zebrafish Proteins/metabolism
  • Tumor Suppressor Proteins/chemistry*
  • Tumor Suppressor Proteins/metabolism
  • Ubiquitin-Specific Proteases/chemistry*
  • Ubiquitin-Specific Proteases/metabolism
  • Signal Transduction
  • Sequence Analysis, Protein
  • Binding Sites
PubMed
25686088 Full text @ Nat. Struct. Mol. Biol.
Abstract
The tumor suppressor CYLD belongs to a ubiquitin (Ub)-specific protease (USP) family and specifically cleaves Met1- and Lys63-linked polyubiquitin chains to suppress inflammatory signaling pathways. Here, we report crystal structures representing the catalytic states of zebrafish CYLD for Met1- and Lys63-linked Ub chains and two distinct precatalytic states for Met1-linked chains. In both catalytic states, the distal Ub is bound to CYLD in a similar manner, and the scissile bond is located close to the catalytic residue, whereas the proximal Ub is bound in a manner specific to Met1- or Lys63-linked chains. Further structure-based mutagenesis experiments support the mechanism by which CYLD specifically cleaves both Met1- and Lys63-linked chains and provide insight into tumor-associated mutations of CYLD. This study provides new structural insight into the mechanisms by which USP family deubiquitinating enzymes recognize and cleave Ub chains with specific linkage types.
Genes / Markers
Figures
Expression
Phenotype
Mutations / Transgenics
Human Disease / Model
Sequence Targeting Reagents
Fish
Antibodies
Orthology
Engineered Foreign Genes
Mapping