PUBLICATION

An EAAT2b/SLC1A2b-mediated chloride leak current enables rapid cone photoreceptor signalling

Authors
Zang, J., Niklaus, S., Neuhauss, S.C.F.
ID
ZDB-PUB-251127-2
Date
2025
Source
Open Biology   15: 250347250347 (Journal)
Registered Authors
Neuhauss, Stephan, Zang, Jingjing
Keywords
anion current, excitatory amino acid transporter, genome duplication, photoreceptor, vision, zebrafish
MeSH Terms
  • Electroretinography
  • Zebrafish/genetics
  • Zebrafish/metabolism
  • Gene Editing
  • Gene Knockout Techniques
  • CRISPR-Cas Systems
  • Retinal Cone Photoreceptor Cells*/metabolism
  • Excitatory Amino Acid Transporter 2*/genetics
  • Excitatory Amino Acid Transporter 2*/metabolism
  • Zebrafish Proteins*/genetics
  • Zebrafish Proteins*/metabolism
  • Animals
  • Signal Transduction*
  • Chlorides*/metabolism
PubMed
41293956 Full text @ Open Biol.
Abstract
Excitatory amino acid transporters not only mediate high-affinity glutamate uptake but also conduct an uncoupled chloride current. In zebrafish, a whole-genome duplication gave rise to two eaat2 paralogues with distinct roles. Excitatory amino acid transporter 2a (SLC1A2b, GLT-1) functions primarily in Müller glia as a glutamate transporter, whereas excitatory amino acid transporter 2b is expressed in cone photoreceptors and exhibits a prominent glutamate-independent chloride current. We hypothesized that this leak current stabilizes the cone resting membrane potential, thereby supporting rapid visual signalling. In order to test this hypothesis, we generated eaat2b knockout zebrafish using CRISPR-Cas9-mediated genome editing. While eaat2b mutants showed no gross morphological abnormalities, they exhibited reduced electroretinogram b-wave amplitudes. Consistent with our hypothesis, eaat2b-deficient larvae displayed a significant reduction in flicker fusion electroretinogram power at each stimulus frequency, indicating impaired temporal processing likely due to delayed repolarization of cone photoreceptors. Our findings reveal a critical role for an excitatory amino acid transporter 2b-mediated chloride anion leak current in regulating the kinetics of photoreceptor responses. This functional innovation, enabled by a whole-genome duplication in the teleost lineage, highlights how gene duplications can lead to the acquisition of physiologically relevant new functions.
Genes / Markers
Figures
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Expression
Phenotype
Mutations / Transgenics
Human Disease / Model
Sequence Targeting Reagents
Fish
Antibodies
Orthology
Engineered Foreign Genes
Mapping