PUBLICATION

FLIM of mCherryTYG Deciphers pH Dynamics and Lifestyles of Salmonella Typhimurium

Authors
Singh, M.K., Fernandez, M., Dilawari, R., Zangoui, P., Kenney, L.J.
ID
ZDB-PUB-250607-5
Date
2025
Source
ACS sensors : (Journal)
Registered Authors
Kenney, Linda
Keywords
Salmonella, bacterial lifestyles, biofilms, fluorescence lifetime imaging, pH, phenotypic heterogeneity, zebrafish, C. elegans
MeSH Terms
  • Hydrogen-Ion Concentration
  • Salmonella typhimurium*/physiology
  • Optical Imaging/methods
  • Biofilms
  • HeLa Cells
  • Humans
  • Animals
  • Microscopy, Fluorescence/methods
  • Fluorescent Dyes*/chemistry
  • Zebrafish
PubMed
40478522 Full text @ ACS Sens
Abstract
Intracellular pH regulation is fundamental to bacterial adaptation, virulence, and survival in diverse environments. Salmonella Typhimurium, a key human pathogen, exploits host and environmental pH cues to transition between planktonic, biofilm, and virulence-associated states. However, precise tools to monitor bacterial pH dynamics at subcellular resolution have been limited. Herein, we report the application of mCherryTYG, a genetically encoded pH-sensitive fluorophore optimized for fluorescence lifetime imaging microscopy (FLIM), enabling robust and high-resolution pH measurements across diverse conditions. mCherryTYG demonstrated exceptional sensitivity across a broad pH range (5.5-8.5) with consistent lifetime responses and was unaffected by temperature, buffer composition, or ionic strength. Using FLIM, we characterized the pH dynamics of Salmonella across in vitro, host, and biofilm contexts. Under acidic stress in vitro, Salmonella maintained a uniform intracellular pH (∼6.04), providing clarity on previously debated heterogeneity. In infections of HeLa cells, Salmonella existed in distinct pH environments: acidic vacuolar pH (∼5.89) and neutral pH (∼7.10). During the late infection stage, ∼17% of the bacterial population retained an acidic pH. Biofilm studies revealed stratified pH profiles with acidic pH near the bottom and neutral pH at the surface, mirroring patterns observed in other pathogens. In heterologous host models, pH gradients shape bacterial adaptation strategies. In C. elegans, Salmonella experienced a progressive internal pH gradient from neutral pH (∼7.10) in the anterior lumen to acidic pH (∼6.45) in the posterior. Similarly, in zebrafish, Salmonella encountered acidic lysosome-rich enterocytes (∼5.84) and neutral regions (∼7.33) in the anterior gut. This study establishes mCherryTYG-FLIM as a transformative tool for studying bacterial pH regulation, revealing pH as a critical modulator of Salmonella lifestyle transitions between virulence and persistence. Our findings provide new insights into host-microbe interactions and present pH as a promising target for therapeutic interventions against bacterial infections.
Genes / Markers
Figures
Expression
Phenotype
Mutations / Transgenics
Human Disease / Model
Sequence Targeting Reagents
Fish
Antibodies
Orthology
Engineered Foreign Genes
Mapping