PUBLICATION

Modular Architecture of the STING C-Terminal Tail Allows Interferon and NF-κB Signaling Adaptation

Authors
de Oliveira Mann, C.C., Orzalli, M.H., King, D.S., Kagan, J.C., Lee, A.S.Y., Kranzusch, P.J.
ID
ZDB-PUB-190425-20
Date
2019
Source
Cell Reports   27: 1165-1175.e5 (Journal)
Registered Authors
Keywords
IRF3, NF-κB, STING, TRAF6, cGAS, innate immunity
MeSH Terms
  • Zebrafish
  • Species Specificity
  • TNF Receptor-Associated Factor 6/genetics
  • TNF Receptor-Associated Factor 6/metabolism*
  • Interferon Type I/metabolism*
  • Protein Conformation
  • Cells, Cultured
  • NF-kappa B/genetics
  • NF-kappa B/metabolism*
  • Mice
  • Humans
  • Animals
  • Immunity, Innate/immunology*
  • Zebrafish Proteins/genetics
  • Zebrafish Proteins/metabolism
  • HEK293 Cells
  • Mice, Knockout
  • Macrophages/immunology*
  • Macrophages/metabolism
  • Membrane Proteins/chemistry
  • Membrane Proteins/genetics
  • Membrane Proteins/metabolism*
  • Membrane Proteins/physiology
  • Interferon Regulatory Factor-3/genetics
  • Interferon Regulatory Factor-3/metabolism*
PubMed
31018131 Full text @ Cell Rep.
Abstract
Stimulator of interferon genes (STING) is a key regulator of type I interferon and pro-inflammatory responses during infection, cellular stress, and cancer. Here, we reveal a mechanism for how STING balances activation of IRF3- and NF-κB-dependent transcription and discover that acquisition of discrete signaling modules in the vertebrate STING C-terminal tail (CTT) shapes downstream immunity. As a defining example, we identify a motif appended to the CTT of zebrafish STING that inverts the typical vertebrate signaling response and results in dramatic NF-κB activation and weak IRF3-interferon signaling. We determine a co-crystal structure that explains how this CTT sequence recruits TRAF6 as a new binding partner and demonstrate that the minimal motif is sufficient to reprogram human STING and immune activation in macrophage cells. Together, our results define the STING CTT as a linear signaling hub that can acquire modular motifs to readily adapt downstream immunity.
Genes / Markers
Figures
Expression
Phenotype
Mutations / Transgenics
Human Disease / Model
Sequence Targeting Reagents
Fish
Antibodies
Orthology
Engineered Foreign Genes
Mapping