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Reprinted from Developmental Biology, 368(2), Johnson, J.L., Hall, T.E., Dyson, J.M., Sonntag, C., Ayers, K., Berger, S., Gautier, P., Mitchell, C., Hollway, G.E., and Currie, P.D., Scube activity is necessary for Hedgehog signal transduction in vivo, 193-202, Copyright (2012) with permission from Elsevier. Full text @ Dev. Biol.